Since your on BES, go to Settings > Options > Security Options > General Settings. Scroll down to Services, press scroll wheel, select Regenerate Encryption Key. After it completes, try to send or reply to a message again.
If that doesn't work, go to Settings > Options > Advanced Options > Service Book. Highlight Desktop[CMIME], press scroll wheel, select Delete. Wait a couple seconds, press scroll wheel, select Undelete. Let the BlackBerry sit for 5-10 minutes while it performs a Slow Sync. You will see a popup message when it completes stating "Activation Complete".
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