The keyboard size is about the same, but the key caps feel a lot better on the 7290 for the vast majority of people. It's much easier to thumb type.
The 7290 quadband also has better reception than the 7730 triband, especially if you go to California, where T-Mobile uses Cingular antennas, and Cingular has 850 Mhz band which the 7730 doesn't have.
Also, at 32megs, you'll see the hourglass less often on the 7290 than the 7730, especially under BlackBerryOS4.
Also, the 7290 benchmarks about 30% faster.
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