If the 8320 / 8820 that you're activating has a GPRS connection, and you don't fill in the third field, it'll activate just like any other Blackberry, over GPRS.
If the BES that you're installing on has had the "WLAN Activation" screen filled in (tick the box and specify the port number to talk to the SMTP server on), and you fill in the third field with an address that the 8320 / 8820 will resolve to the Blackberry Router (or BES, if the router's on the BES), then activation can occur over WiFi without the ETP message being sent all the way to RIM via carrier GPRS.
To test this out, turn on an 8320 / 8820 and turn off GPRS, leaving WiFi on (and configured for the local WLAN of course). Provided that the device can talk to the Router and the ETP message can get to the BES, you can do an activation without GPRS being enabled.
The RIM documentation also implies to me that you can enterprise activate a WiFi Blackberry on one BES using an entirely unconnected Blackberry Router, although I've not tried that though and may be reading WAY too much into the documentation).
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